This
information courtesy of the Colorado Natural Areas Program
Family:
Scrophulariaceae (Figwort)
Other Names:
broad-leaved
toadflax, wild snapdragon
USDA Code:
LIDAM
Legal Status:
Colorado Noxious
Weed List B
Identification
Growth form:
Perennial forb
Flower:
Flowers are borne in
loose, elongate, terminal racemes. Flowers are bright yellow
and resemble snapdragons.
Seeds/Fruit:
Fruits are
egg-shaped to nearly round capsules. Seeds are sharply
angular, and slightly winged.
Leaves:
Leaves are broad,
ovate to ovate-lanceolate, and are alternate, generally
clasping but crowded.
Stems:
Mature plants are up
to three feet tall. A single toadflax plant contains from
1-25 vertical, floral stems. which are thick-walled and
somewhat woody.
Roots:
The taproot may
penetrate one meter into the soil. Horizontal roots may grow
to be several meters long, and can develop adventitious buds
that may form independent plants.
Seedling:
No information
available.
Similar
Species
Exotics:
Yellow toadflax (Linaria
vulgaris) is similar in appearance, but has more linear
pointed leaves, and is generally a smaller plant.
Natives:
None known.
Impacts
Agricultural:
Low-till
cultivation practices have contributed to the resurgence of
toadflax populations on agricultural lands (McClay 1992).
Dalmatian toadflax contains a glucoside, a quinoline
alkaloid, and peganine which make it toxic to livestock
(Rees et al. 1996). However, Dalmatian toadflax is
generally considered unpalatable, and reports of livestock
poisonings are rare.
Ecological:
Dalmatian
toadflax is a persistent, aggressive invader and capable of
forming colonies through adventitious buds from creeping
root systems. These colonies can push out native grasses and
other perennials, thereby altering the species composition
of natural communities. New infestations of Dalmatian
toadflax can occur in naturally occurring disturbances or in
small openings in pristine or excellent-condition rangeland
(Lajeunesse 1999). Dalmatian toadflax can rapidly
colonize open sites. It is most commonly found along
roadsides, fences, rangelands, croplands, clear cuts, and
pastures. Disturbed or cultivated ground is a prime
candidate for colonization. Toadflax can significantly
reduce crop yields and stress native communities. In one
study, toadflax-free plots produced 2.5 times more grass
than plots where toadflax was present (Robocker 1974). The
seedlings of toadflax are considered ineffective competitors
for soil moisture with established perennials and winter
annuals (Morishita 1991). However, once established both
species of toadflax suppress other vegetation mainly by
intense competition for limited soil water. Mature plants
are particularly competitive with winter annuals and
shallow-rooted perennials (Robocker 1974).
Human:
No information
available.
Habitat
and Distribution
General requirements:
Dalmatian
toadflax can adapt its growth to fit a wide range of
environmental conditions, and is tolerant of low
temperatures and coarse-textured soils.
Distribution:
In Colorado,
Dalmatian toadflax is commonly found between 5,000 to
6,500 feet in oak, aspen, sagebrush, mountain brush, and
riparian communities. It can be found growing up to 9,000
feet (A. Green)
Historical:
Native to
Mediterranean region.
Biology/Ecology
Life cycle:
Spring emergence
occurs about mid-April and depends primarily on temperature.
During the first year the plant forms a rosette and develops
a deep root system. Prostrate stems emerge in September and
produce ovate leaves. Prostrate stems are tolerant to
freezing and are associated with floral stem production the
following year (Robocker 1974). The strong upright floral
stems that characterize mature toadflax plants develop after
a winter’s dormancy, and emerge about the same time as new
seedlings in mid-April. A single plant will produce from
1-25 floral stems. Flowering occurs from May-August and
seeds mature from July-September. Dalmatian toadflax can
also reproduce vegetatively. Stems develop from adventitious
buds on primary and lateral roots. Vegetative reproduction
from root buds can occur as early as 2-3 weeks after
germination, and is possible from root fragments as short as
1 cm in length (Zimmerman 1996). These buds can grow their
own root and shoot systems, and become independent plants
the next year. In addition to promoting growth, the large,
deep, root systems of Dalmatian toadflax exploit water
efficiently. The taproot may penetrate 3-4 feet into the
soil and lateral roots may be 6-12 feet long. (periods of
growth, emergence, bolting, flowering and seed production)
Mode of reproduction:
By
seeds and vegetatively
Seed production:
A mature
Dalmatian toadflax can produce up to 500,000 seeds annually
(Morishita 1991).
Seed bank:
Seeds may remain
viable in the soil for up to ten years.
Dispersal:
No information
available.
Integrated
Management Summary
Management of Dalmatian toadflax
must focus on both reducing the rate of vegetative spread
and reducing seed production (Lajeunesse 1999). Successful
management requires integrating as many control tactics as
possible. Dalmatian toadflax has high genetic variability,
and local populations can respond differently to control
actions, especially herbicide treatments. Successful control
can be obtained by pulling, or killing the plants with
herbicide before toadflax seed production begins (Carpenter
and Murray 1998). Since the plant also spreads through
vegetative propagation, and the seeds can remain dormant for
up to ten years, this process must be repeated every year
for at least ten years to completely remove a stand.
Competitive perennial grasses and forbs should be planted to
utilize water and nutrients that would otherwise be readily
available to toadflax.
CONTROL
CSU
Fact Sheet
References
Carpenter, A.T. and T.A. Murray.
1998. Element Stewardship Abstract for Linaria dalmatica.
The Nature Conservancy, Wildland Weed Program. Internet:
1/21/99. Available: http://tncweeds.ucdavis.edu/esadocs.html
Lajeunesse, S. 1999. Dalmatian
and yellow toadflax. In: R.L. Sheley and J.K. Petroff (eds.)
Biology and management of noxious rangeland weeds. Oregon
State University Press, Corvallis. pp. 202-216.
McClay,
A.S. 1992. Effects of Brachypterolus
pulicarius (L.) (Coleoptera: Nitidulidae) on flowering
and seed production of common toadflax. The Canadian
Entomologist 124: 631-636.
Moroshita,
D.W. 1991. Dalmatian
toadflax, yellow toadflax, black henbane, and tansymustard:
importance, distribution, and control. Pages 399-408 In:
L.F. James, J.O. Evans, M.H. Ralphs, and R.D. Child
(editors) Noxious range weeds. Westview Press, Boulder, San
Francisco, and Oxford.
Robocker,
W.C. 1974. Life
history, ecology, and control of Dalmatian toadflax. Tech.
Bull No. 79. Wash. Agric. Exp. Stn.
Zimmerman, J.A.C. 1996. Ecology
and distribution of Linaria vulgaris (L.) Miller,
Scrophulariaceae. USGS Biological Resources Division,
Colorado Plateau Field Station-Flagstaff, Arizona. Internet.
01/14/98. Available: http://www.nbs.nau.edu/FNF/Vegetation/Exotics/linariatitle.html
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