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Cheatgrass
Downy
brome
Bromus
tectorum (L.); Anisantha tectorum (L.) Nevski
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Keys
to Identification:
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Family:
Poaceae (Grass)
Other Names: cheatgrass,
downy chess, early chess, drooping brome, downy cheat,
slender chess, downy bromegrass, military grass, broncograss,
Mormon oats
USDA Code: BRTE,
ANTE6
Legal Status: Colorado
List C
Identification
Growth form: Annual
or winter annual grass
Flower: Downy
brome panicles (loose, irregularly compound flowering part
of plant with flowers borne on individual stalks) change
color from green to purple to brown as the plant matures and
eventually dries out. Branches are slender, drooping, hairy,
flexuous, with up to eight spikelets.
Seeds/Fruit: Spikelets
including awns are 0.8-2 inches long, nodding, with 2-8
florets.
Leaves: Leaves
are light-green and hairy. Sheaths are fused except near the
node at the bottom of each sheath. The lower sheaths are
conspicuously hairy, while the upper sheaths are sometimes
smooth.
Stems: Mature
plants are generally 4-30 inches tall. Stems are
erect, slender, glabrous, or slightly hairy.
Roots: Finely
divided fibrous root system that can reach a depth of about
12 inches.
Seedling: Downy
brome seedlings can usually be distinguished by their hairy
leaf blades and sheaths.
Similar Species
Exotics: Similar
to Japanese brome (Bromus japonicus).
Natives: None
known.
Impacts
Ecological: Downy
Brome can greatly alter the species composition of dry
native rangeland vegetation by competitive exclusion of
reproduction of native plant species and by facilitation of
wildfires (Mosky et al. 1999). Invasion of downy
brome is greatest in drier environments, particularly in
sagebrush-steppe communities. The process in which a
pristine Great Basin shrub-steppe ecosystem deteriorates
into one that is dominated by downy brome takes several
years and has several distinct cycles. First, some sort of
disturbance, such as heavy grazing, allows downy brome and
other annuals to invade and proliferate. The dry stands of
downy brome in the summer increase the occurrence of
frequent fires. Initially, this creates an environment
dominated by annual grasses, matchweed (Gutierrezia
sarothrae), and rabbitbrush (Chrysothamnus spp.).
As fires become even more frequent, the area will be
dominated by annual grasses alone, with the loss of surface
soil, nutrients, and near permanent deterioration of the
site. Species that are commonly displaced by downy brome
include big sagebrush (Artemisia tridentata),
antelope bitterbrush (Purshia tridentata), bluebunch
wheatgrass (Agropyron spicatum = Pseudoroegneria
spicata ), crested wheatgrass (Agropyron
cristatum), western wheatgrass (Agropyron
smithii = Pascopyrum smithii), Sandberg bluegrass
(Poa sandbergii = Poa secunda),
needle-and-thread grass (Stipa comata =
Hesperostipa comata), and Thurber’s needlegrass (Stipa
thurberiana)
Habitat and
Distribution
General
requirements: Downy brome is common in recently burned
rangeland and wildlands, winter crops, waste areas,
abandoned fields, eroded areas, and overgrazed grasslands (Upadhyaya
et al. 1986). It can invade rangelands that have never been
grazed by livestock (Svejcar and Tausch 1991).
Distribution:
Common throughout Colorado from 4,000 to 9,000 feet. Widely
distributed throughout North America. Although downy brome
occurs in a variety of habitats, it is most prominent on the
Columbia-Snake River Plateau, Wyoming Basin, and the
northern edge of the Great Basin in disturbed sagebrush
steppe communities (Rice and Mack 1991, West 1983).
Biology/Ecology
Life cycle: Vast
numbers of downy brome seedlings usually germinate after the
first fall rain in infested areas (West 1983). The leaves
typically grow little in the fall, and plants are normally
1-2 inches in height when covered by snow in December. The
young, fall-germinated seedlings often over-winter in a
semi-dormant state and complete their lifecycle the
following spring (Upadhyaya et al. 1986). However, downy
brome roots can grow in soil temperatures approaching
freezing (West 1983), and roots will continue to grow
throughout the winter until soil temperatures drop below
about 37 degrees F. Plants head in late April to early May
followed by flowering within a week (Upadhyaya et al. 1986).
The seeds reach the soft dough-stage in mid to late May, and
mature in mid to late June (Upadhyaya et al. 1986).
During ripening
downy flowers turn purple and then brown as they mature.
Once the seeds have matured, downy brome plants dry and
become flammable. There is a correlation between plant color
and moisture status during the drying process (FEIS 1998).
Downy brome passes from green (>100% moisture content),
to a purple hue (30-100% moisture content), to a straw color
(<30% moisture content) as it dries (FEIS 1998). The
onset of purple coloring should be taken as a warning that
hazardous fire conditions will develop within two weeks (FEIS
1998).
Downy brome greens
up earlier in the spring than most other species. Depletion
of soil moisture is a mechanism by which downy brome
suppresses seedlings of desirable, perennial grasses (Melgoza
et al. 1990). In addition, thick mulch in dense
cheatgrass stands favors cheatgrass seedling germination and
establishment while inhibiting that of perennial
bunchgrasses (Svejar and Tausch 1991).
Mode of
reproduction: Reproduces by seeds.
Seeds production: Downy
brome can be a prolific seed producer and is capable of
producing up to 400 lbs. of seeds acre (Upadhyaya et al.
1986). Seed production per culm, per plant, and per unit
area is dependent on plant density and environmental factors
(Upadhyaya et al. 1986). Average seed production per plant
is generally lowest where plant density is highest (Rice and
Mack 1991). If precipitation is adequate, the majority of
downy brome seeds will germinate in the fall, or within a
year of maturation (Upadhyaya et al. 1986). However, dry
conditions can cause environmentally induced dormancy, which
may last several years and break down at erratic intervals
(Young and Evans 1985).
Seed bank: Low
survival rates in the soil, but seeds may last in the seed
bank for a few years.
Dispersal: Seeds
are dispersed short distances by wind, and the long awns can
attach to the fur or feathers of an animal, as well as
clothing.
Integrated
Management Summary
Downy brome is
characterized by its early maturation, high seed production,
and vigorous growth under favorable environmental
conditions. Although, downy brome readily invades perennial
forage crops and rangeland under poor management, it also
invades communities in the absence of disturbance (Douglas
et al. 1990). In undisturbed sites, downy brome will most
commonly spread along soil cracks and work its way outward
into the natural community (Rice and Mack 1991). Downy
brome has a dual role as a serious weed and important early
season forage for cattle and sheep (Upadhyaya et al. 1986).
Downy brome provides the bulk of early spring forage for all
classes of stock on grazing lands in the Intermountain and
Pacific Northwest regions (Upadhyaya et al. 1986). Lasting
control of downy brome will require a combination of
chemical control, physical control, vegetative suppression,
and proper livestock management where land is grazed. This
"cumulative stress" method will keep the plants
constantly under stress, reducing their ability to flourish
and spread. Also, a cumulative stress approach provides a
level of redundancy in case one type of treatment is not
implemented or proves to be ineffective (Carpenter and
Murray 1998).
References
Carpenter, A.T.
and T.A. Murray. 1998. Element Stewardship Abstract for Bromus
tectorum. The Nature Conservancy, Wildland Weed
Program. Internet: http://tncweeds.ucdavis.edu
Douglas, B.J.,
A.G. Thomas, and D.A. Derksen. 1990. Downy brome (Bromus
tectorum) invasion into southwestern Saskatchewan.
Canadian Journal of Plant Science 70:1143-1151.
FEIS - Fire
Effects Information System [Online] (1996, September).
Prescribed Fire and Fire Effects Research Work Unit, Rocky
Mountain Research Station (producer), US Forest Service.
Available: www.fs.fed.us/database/feis/ [1998,March 12]
Melgoza, G., R.S.
Nowak and R.J. Tausch. 1990. Soilwater exploration after
fire: competition between Bromus tectorum and two
native species. Oceologia 83:7-13.
Mosely, J.C., S.C.
Bunting, and M.E. Monoukian. 1999. Cheatgrass. In: R.L.
Sheley and J.K. Petroff (eds.) Biology and management of
noxious rangeland weeds. Oregon State University Press,
Corvallis. pg. 175-188.
Mosely, J.C. 1996.
Prescribed sheep grazing to suppress cheatgrass: A review.
Sheep and Goat Research Journal 12:74-81.
Ponzetti, J.M.
1997. Assessment of medusahead and cheatgrass control
techniques at Lawrence Memorial Grassland Preserve 1996
Annual Report. The Nature Conservancy of Oregon. January
14, 1997.
Rice, K.J., and
R.N. Mack. 1991. Ecological genetics of Bromus tectorum:
intraspecific variation in phenotypic plasticity.
Oecologia 88:84-90.
Svejcar, T. and R.
Tausch. 1991. Anako Island, Nevada: A relict area
dominated by annual invader species. Rangelands 13:
233-236.
Upadhaya, M.K., R.
Turkington, and D. McIlvride. 1986. The biology of
Canadian weeds. 75. Bromus tectorum L. Canadian
Journal of Plant Science 66:689-709.
West, N.E. 1983.
Western intermountain sagebrush steppe. Temperate
Deserts and Semi-Deserts. Elsevier Scientific
Publishing Company, Amsterdam. pp. 351-373.
Whitson, T.D. and
D.W. Koch. 1998. Control of downy brome (Bromus
tectorum) with herbicides and perennial grass
competition. Weed Technology 12:391-396.
Wiese, A.F., C.D.
Salisbury, and B.W. Bean. 1995. Downy brome (Bromus
tectorum), jointed goatgrass (Aegilops cylindrica)
and horseweed (Conyza canadensis) control in
fallow. Weed Technology 9:249-254.
Young, J.A. and
R.A. Evans. 1985. Demography of Bromus tectorum in Artemisia
communities. J. White (ed.), The Population Structure
of Vegetation. Dr. W. Junk Publishers, Dordrecht,
Netherlands.
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